diff --git a/pw_docgen/py/pw_docgen/seed.py b/pw_docgen/py/pw_docgen/seed.py
index 063f2d6..0b1b5ee 100644
--- a/pw_docgen/py/pw_docgen/seed.py
+++ b/pw_docgen/py/pw_docgen/seed.py
@@ -78,17 +78,17 @@
 
             if 'changelist' in meta:
                 # The SEED has not yet been merged and points to an active CL.
-                change_url = (
+                change_link = (
                     'https://pigweed-review.googlesource.com'
                     f'/c/pigweed/pigweed/+/{meta["changelist"]}'
                 )
-                title = f'`{meta["title"]} <{change_url}>`__'
+                title = f'`{meta["title"]} <{change_link}>`__'
                 seed_toctree.append(
-                    f'{meta["number"]}: {meta["title"]}<{change_url}>',
+                    f'{meta["number"]}: {meta["title"]}<{change_link}>',
                 )
             else:
                 # The SEED document is in the source tree.
-                title = f':ref:`seed-{meta["number"]}`'
+                title = f':ref:`{meta["title"]} <seed-{meta["number"]}>`'
                 seed_toctree.append(Path(meta["rst_file"]).stem)
 
             seed_table.extend(
diff --git a/seed/0001-the-seed-process.rst b/seed/0001-the-seed-process.rst
index 0cde0e5..c99fbee 100644
--- a/seed/0001-the-seed-process.rst
+++ b/seed/0001-the-seed-process.rst
@@ -183,6 +183,7 @@
 
       pw_seed("5309") {
         sources = [ "5309-pw_rpc-over-smoke-signals.rst" ]
+        title = "pw_rpc Over Smoke Signals"
         status = "Accepted"
         author = "Your Name"
       }
diff --git a/seed/BUILD.gn b/seed/BUILD.gn
index b9e72de..7347f5e 100644
--- a/seed/BUILD.gn
+++ b/seed/BUILD.gn
@@ -19,24 +19,28 @@
 pw_seed("0001") {
   sources = [ "0001-the-seed-process.rst" ]
   inputs = [ "0001-the-seed-process/seed-index-gerrit.png" ]
+  title = "The SEED Process"
   status = "Meta"
   author = "The Pigweed Authors"
 }
 
 pw_seed("0002") {
   sources = [ "0002-template.rst" ]
+  title = "SEED Template"
   status = "Meta"
   author = "The Pigweed Authors"
 }
 
 pw_seed("0101") {
   sources = [ "0101-pigweed.json.rst" ]
+  title = "pigweed.json"
   status = "Accepted"
   author = "Rob Mohr"
 }
 
 pw_seed("0102") {
   sources = [ "0102-module-docs.rst" ]
+  title = "Consistent Module Documentation"
   status = "Accepted"
   author = "Chad Norvell"
 }
@@ -50,12 +54,14 @@
 
 pw_seed("0104") {
   sources = [ "0104-display-support.rst" ]
+  title = "Display Support"
   status = "Accepted"
   author = "Chris Mumford"
 }
 
 pw_seed("0105") {
   sources = [ "0105-pw_tokenizer-pw_log-nested-tokens.rst" ]
+  title = "Nested Tokens and Tokenized Log Arguments"
   status = "Accepted"
   author = "Gwyneth Chen"
 }
@@ -69,48 +75,56 @@
 
 pw_seed("0107") {
   sources = [ "0107-communications.rst" ]
+  title = "Pigweed Communications"
   status = "Accepted"
   author = "Wyatt Hepler"
 }
 
 pw_seed("0108") {
   sources = [ "0108-pw_emu-emulators-frontend.rst" ]
+  title = "Emulators Frontend"
   status = "Accepted"
   author = "Octavian Purdila"
 }
 
 pw_seed("0109") {
   sources = [ "0109-comms-buffers.rst" ]
+  title = "Communication Buffers"
   status = "Accepted"
   author = "Taylor Cramer"
 }
 
 pw_seed("0110") {
-  status = "Accepted"
   sources = [ "0110-memory-allocation-interfaces.rst" ]
+  title = "Memory Allocation Interfaces"
+  status = "Accepted"
   author = "Alexei Frolov"
 }
 
 pw_seed("0111") {
   sources = [ "0111-build-systems.rst" ]
+  title = "Make Bazel Pigweed's Primary Build System"
   status = "Accepted"
   author = "Ted Pudlik"
 }
 
 pw_seed("0112") {
   sources = [ "0112-async-poll.rst" ]
+  title = "Async Poll Model"
   status = "Accepted"
   author = "Taylor Cramer"
 }
 
 pw_seed("0113") {
   sources = [ "0113-bazel-cc-toolchain-api.rst" ]
+  title = "Add modular Bazel C/C++ toolchain API"
   status = "Accepted"
   author = "Armando Montanez"
 }
 
 pw_seed("0114") {
   sources = [ "0114-channels.rst" ]
+  title = "Channels"
   status = "Accepted"
   author = "Wyatt Hepler"
 }
diff --git a/seed/seed.gni b/seed/seed.gni
index cbbfc04..134dacf 100644
--- a/seed/seed.gni
+++ b/seed/seed.gni
@@ -74,7 +74,7 @@
 #  changelist (optional): If the SEED has not yet been merged, the number of the
 #      CL on Gerrit where the SEED is being reviewed.
 #
-#  title (optional): If `changelist` is provided, the title of the SEED.
+#  title (required): The title of the SEED.
 #
 #  authors (required): Comma-separated list of SEED authors.
 #
@@ -87,6 +87,7 @@
   _has_sources = defined(invoker.sources) && invoker.sources != []
   assert(_has_sources || defined(invoker.changelist),
          "A SEED must either have in-tree sources or an active CL")
+  assert(defined(invoker.title), "SEEDs must have a title")
   assert(defined(invoker.status), "SEEDs must list their status")
   assert(defined(invoker.author), "SEEDs must list their author(s)")
 
@@ -95,6 +96,7 @@
 
   _metadata = {
     number = target_name
+    title = invoker.title
     status = invoker.status
     author = invoker.author
   }
@@ -103,10 +105,7 @@
     _sources = invoker.sources
     _metadata.rst_file = _sources[0]
   } else {
-    assert(defined(invoker.title),
-           "SEEDs pointing to a CL must provide a title")
     _metadata.changelist = invoker.changelist
-    _metadata.title = invoker.title
   }
 
   generated_file(_metadata_target) {